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"The researchers suspect that the growth of these hollow structures is tied to the specific physical and biochemical conditions near the halocline, the layer that separates the freshwater from the underlying saltwater. "Microbes involved in the nitrogen cycle, which are still active today, could have played a major role in calcite precipitation because of their ability to increase the pH," explains Dr Stinnesbeck."
https://www.sciencedaily.com/releases/2017/11/171124114942.htm